Selected Quarterly Financial Data (Unaudited) (Tables)

12 Months Ended

Dec. 31, 2016

Selected Quarterly Financial Information [Abstract]

Schedule of Selected Quarterly Results from Operations

Selected quarterly results from operations for the years ended December 31, 2016 and 2015 are as follows (in thousands, except per share amounts):

	2016 Quarter Ended
	March 31,			June 30,			September30,			December31,
Grant revenues	$	859		$	1,373		$	1,149		$	1,247
Loss from operations		(4,567	)		(5,495	)		(6,301	)		(5,543	)
Net loss		(4,547	)		(5,430	)		(6,238	)		(5,483	)
Basic and diluted net loss per common share	$	(7.10	)	$	(0.46	)	$	(0.47	)	$	(0.41	)

	2015 Quarter Ended
	March 31,			June 30,			September30,			December31,
Grant revenues	$	—		$	3,427		$	1,073		$	1,585
Loss from operations		(2,469	)		(1,432	)		(3,417	)		(3,997	)
Net loss		(2,468	)		(1,427	)		(3,413	)		(3,987	)
Basic and diluted net loss per common share	$	(4.71	)	$	(2.44	)	$	(5.57	)	$	(6.36	)