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License and Collaboration Agreements (Details) - USD ($)
1 Months Ended 12 Months Ended
Feb. 28, 2015
Dec. 31, 2019
Dec. 31, 2015
Feb. 28, 2017
Monash License        
License and Collaboration Agreements        
Minimum annual fee   $ 25,000    
Aggregate potential milestone payments   $ 45,100,000    
Period of written notice to terminate   30 days    
Monash License | Research and development        
License and Collaboration Agreements        
One-time cash payment   $ 275,000    
Licensing Agreement | Scripps Research Institute        
License and Collaboration Agreements        
Minimum annual fee     $ 25,000  
Aggregate potential milestone payments     $ 2,600,000  
Milestone payments made   $ 0    
Period of written notice to terminate     90 days  
Licensing Agreement | Scripps Research Institute | Research and development        
License and Collaboration Agreements        
One-time cash payment     $ 10,000  
Licensing Agreement | Vernalis (R&D) Limited        
License and Collaboration Agreements        
Milestone payments made       $ 3,000,000
Period of written notice to terminate 90 days      
Licensing Agreement | Vernalis (R&D) Limited | Minimum        
License and Collaboration Agreements        
Aggregate potential milestone payments $ 220,000,000      
Licensing Agreement | Vernalis (R&D) Limited | Research and development        
License and Collaboration Agreements        
One-time cash payment $ 1,000,000