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Revenue - Collaboration Agreement - Description of good or service to be transferred in contract with customer (Details) - Astellas Collaboration Agreement
$ in Millions
3 Months Ended
Jan. 13, 2020
USD ($)
item
Mar. 31, 2020
item
Collaboration and License Agreement    
Upfront payment received | $ $ 50.0  
Number of co-development targets 3 3
Number of independent targets 2 2