X0202 SCHEDULE 13G 0001716774 XXXXXXXX LIVE Ordinary Shares, nominal value EUR0.04 per share 06/30/2026 0001612940 ProQR Therapeutics NV N71542109 Zernikedreef 9 Leiden The Netherlands XX CK 2333 Rule 13d-1(b) Aberdeen Group plc X0 0 8932507 0 8932507 8932507 N 6.33 HC abrdn Inc. X1 0 8932507 0 8932507 8932507 N 6.33 CO IA ProQR Therapeutics NV Zernikedreef 9, Leiden, The Netherlands CK 2333 Aberdeen Group plc abrdn Inc. Aberdeen Group plc: 1 George Street Edinburgh, United Kingdom EH2 2LL abrdn Inc.: 1900 Market Street, Suite 200 Philadelphia PA 19103 Aberdeen Group plc - UNITED KINGDOM abrdn Inc. - UNITED STATES N HC 8,932,507 6.33 Aberdeen Group plc - 0 abrdn Inc. - 0 Aberdeen Group plc - 8,932,507 abrdn Inc. - 8,932,507 Aberdeen Group plc - 0 abrdn Inc. - 0 Aberdeen Group plc - 8,932,507 abrdn Inc. - 8,932,507 Y Y N See Exhibit Y Y N By signing below I certify that, to the best of my knowledge and belief, the securities referred to above were acquired and are held in the ordinary course of business and were not acquired and are not held for the purpose of or with the effect of changing or influencing the control of the issuer of the securities and were not acquired and are not held in connection with or as a participant in any transaction having that purpose or effect, other than activities solely in connection with a nomination under § 240.14a-11. Aberdeen Group plc abrdn Holdings Limited abrdn Inc. Aberdeen Group plc is the parent company. abrdn Holdings Limited is the intermediate holding company for abrdn Inc. abrdn Inc. beneficially owns on behalf of our underlying clients 5% or greater of the outstanding shares of the security class being reported on this Schedule 13G. Aberdeen Group plc R. Chaudhry 07/29/2026 abrdn Inc. R. Chaudhry 07/29/2026