Collaboration Revenue (Tables)

3 Months Ended

Mar. 31, 2020

Revenue From Contract With Customer [Abstract]

Changes in Contract Liabilities

The following table presents changes in the Company’s contract liabilities during the three months ended March 31, 2020 and 2019:

	Balance as of December 31, 2019		Additions		Deductions			Balance as of March 31, 2020
Three months ended March 31, 2020
Contract liabilities:
Deferred revenue - related party		110,071		—		(5,462	)		104,609
Deferred revenue		9,668		816		(1,988	)		8,496

	Balance as of December 31, 2018		Additions		Deductions			Balance as of March 31, 2019
Three months ended March 31, 2019
Contract liabilities:
Deferred revenue - related party		137,259		—		(6,615	)		130,644
Deferred revenue		—		6,667		(260	)		6,407

During the three months ended March 31, 2020 and 2019 the Company recognized the following revenues as a result of changes in the contract liabilities balances in the respective periods (in thousands):

	Three Months Ended March 31,
	2020		2019
Revenue recognized in the period from:
Amounts included in the contract liability at the beginning    of the period		6,634		6,615