Condensed Consolidated Statements of Operations and Comprehensive Loss - USD ($) $ in Thousands	3 Months Ended		6 Months Ended
	Jun. 30, 2022	Jun. 30, 2021	Jun. 30, 2022	Jun. 30, 2021
Revenue Not from Contract with Customer	$ 207		$ 207
Revenues, Total	$ 3,215	$ 4,539	$ 6,084	$ 9,993
Revenue, Product and Service [Extensible Enumeration]	Collaboration [Member]	Collaboration [Member]	Collaboration [Member]	Collaboration [Member]
Operating expenses:
Research and development	$ 18,760	$ 17,682	$ 35,771	$ 32,422
General and administrative	6,958	5,933	13,870	12,054
Total operating expenses	25,718	23,615	49,641	44,476
Loss from operations	(22,503)	(19,076)	(43,557)	(34,483)
Other income (expense):
Interest income	156	11	172	20
Other income (expense), net		(5)	111	(6)
Total other income, net	267	6	283	14
Net loss and comprehensive loss	$ (22,236)	$ (19,070)	$ (43,274)	$ (34,469)
Net loss per share, basic	$ (0.78)	$ (0.68)	$ (1.53)	$ (1.27)
Net loss per share, diluted	$ (0.78)	$ (0.68)	$ (1.53)	$ (1.27)
Weighted-average common shares outstanding, basic	28,367,355	27,919,647	28,256,810	27,100,817
Weighted-average common shares outstanding, diluted	28,367,355	27,919,647	28,256,810	27,100,817
Collaboration [Member]
Revenue	$ 3,008	$ 4,539	$ 5,877	$ 9,993