Condensed Consolidated Statements of Operations and Comprehensive Loss - USD ($) $ in Thousands	3 Months Ended
	Mar. 31, 2022	Mar. 31, 2021
Total revenue	$ 2,869	$ 5,454
Revenue, Product and Service [Extensible Enumeration]	Collaboration [Member]	Collaboration [Member]
Operating expenses:
Research and development	$ 17,010	$ 14,740
General and administrative	6,912	6,120
Total operating expenses	23,922	20,860
Loss from operations	(21,053)	(15,406)
Other income (expense):
Interest income	15	9
Other income (expense), net		(2)
Total other income, net	15	7
Net loss and comprehensive loss	$ (21,038)	$ (15,399)
Net loss per share, basic and diluted	$ (0.75)	$ (0.59)
Weighted-average common shares outstanding, basic and diluted	28,145,036	26,264,019