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Selected Quarterly Financial Information (Unaudited) (Tables)
 12 Months Ended
Dec. 31, 2017
Quarterly Financial Information Disclosure [Abstract]  
Quarterly Financial Information

 

 

 

Quarters Ended

 

 

 

March 31, 2017

 

 

June 30, 2017

 

 

September 30, 2017

 

 

December 31, 2017

 

Total revenues

 

$

455

 

 

$

6,562

 

 

$

1,336

 

 

$

2,040

 

Research and development expense

 

$

16,619

 

 

$

13,917

 

 

$

12,518

 

 

$

14,170

 

General and administrative expense

 

$

6,622

 

 

$

6,355

 

 

$

9,444

 

 

$

4,808

 

Total operating expenses

 

$

23,377

 

 

$

21,618

 

 

$

22,645

 

 

$

18,638

 

Net loss

 

$

(21,993

)

 

$

(14,473

)

 

$

(20,706

)

 

$

(15,997

)

Basic and diluted net loss per common share

 

$

(0.82

)

 

$

(0.47

)

 

$

(0.67

)

 

$

(0.51

)

 

 

 

Quarters Ended

 

 

 

March 31, 2016

 

 

June 30, 2016

 

 

September 30, 2016

 

 

December 31, 2016

 

Total revenues

 

$

393

 

 

$

2,375

 

 

$

125

 

 

$

1,696

 

Research and development expense

 

$

6,183

 

 

$

10,680

 

 

$

12,560

 

 

$

16,059

 

General and administrative expense

 

$

5,479

 

 

$

6,169

 

 

$

6,200

 

 

$

5,742

 

Total operating expenses

 

$

11,644

 

 

$

17,327

 

 

$

18,793

 

 

$

22,165

 

Net loss

 

$

(10,768

)

 

$

(14,437

)

 

$

(18,154

)

 

$

(19,608

)

Basic and diluted net loss per common share

 

$

(0.41

)

 

$

(0.55

)

 

$

(0.69

)

 

$

(0.74

)