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Selected Quarterly Financial Information (Unaudited) (Tables)
 12 Months Ended
Dec. 31, 2016
Quarterly Financial Information Disclosure [Abstract]  
Quarterly Financial Information

 

 

 

Quarters Ended

 

 

 

March 31, 2016

 

 

June 30, 2016

 

 

September 30, 2016

 

 

December 31, 2016

 

Total revenues

 

$

393

 

 

$

2,375

 

 

$

125

 

 

$

1,696

 

Research and development expense

 

$

6,183

 

 

$

10,680

 

 

$

12,560

 

 

$

16,059

 

General and administrative expense

 

$

5,479

 

 

$

6,169

 

 

$

6,200

 

 

$

5,742

 

Total operating expenses

 

$

11,644

 

 

$

17,327

 

 

$

18,793

 

 

$

22,165

 

Net loss

 

$

(10,768

)

 

$

(14,437

)

 

$

(18,154

)

 

$

(19,608

)

Basic and diluted net loss per common share

 

$

(0.41

)

 

$

(0.55

)

 

$

(0.69

)

 

$

(0.74

)

 

 

 

Quarters Ended

 

 

 

March 31, 2015

 

 

June 30, 2015

 

 

September 30, 2015

 

 

December 31, 2015

 

Total revenues

 

$

644

 

 

$

1,363

 

 

$

1,140

 

 

$

4,441

 

Research and development expense

 

$

2,791

 

 

$

4,012

 

 

$

5,664

 

 

$

4,812

 

General and administrative expense

 

$

1,716

 

 

$

3,397

 

 

$

2,567

 

 

$

4,232

 

Total operating expenses

 

$

4,637

 

 

$

7,659

 

 

$

8,487

 

 

$

9,942

 

Net loss

 

$

(4,011

)

 

$

(6,290

)

 

$

(7,332

)

 

$

(5,178

)

Basic and diluted net loss per common share

 

$

(0.94

)

 

$

(3.24

)

 

$

(1.52

)

 

$

(0.20

)