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Significant Agreements (Tables)
 12 Months Ended
Dec. 31, 2016
Organization Consolidation And Presentation Of Financial Statements [Abstract]  
Schedule of License Revenue

License revenue consists of the following (in thousands):

 

 

 

Years Ended December 31,

 

 

 

2016

 

 

2015

 

 

2014

 

Up-front fees and option fees for commercial licenses

 

$

3,050

 

 

$

5,900

 

 

$

4,000

 

Maintenance fees for commercial licenses

 

 

910

 

 

 

635

 

 

 

385

 

Milestone fees

 

 

—

 

 

 

250

 

 

 

—

 

Research and other license revenue

 

 

343

 

 

 

240

 

 

 

190

 

 

 

$

4,303

 

 

$

7,025

 

 

$

4,575