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SHARE-BASED COMPENSATION (Tables)
 3 Months Ended
Mar. 31, 2020
Share-based Payment Arrangement [Abstract]  
Stock Option Activity
A summary of Common Stock option activity is as follows:
 
Number of Options
(in thousands)
 
Weighted Average Exercise Price
 
Weighted Average Remaining Contractual Term
(years)
 
Aggregate Intrinsic Value
(in thousands)
Outstanding at December 31, 2019
3,497

 
$
5.19

 
5.91
 
$
—

Granted
1,778

 
$
0.89

 
 
 
 
Exercised
(17
)
 
$
0.76

 
 
 
 
Forfeited or expired
(453
)
 
$
6.62

 
 
 
 
Outstanding at March 31, 2020
4,805

 
$
3.48

 
6.93
 
$
—

Vested or expected to vest at March 31, 2020
4,805

 
$
3.48

 
6.93
 
$
—

Exercisable as of March 31, 2020
2,099

 
$
4.84

 
5.42
 
$
—

Vested as of March 31, 2020
2,099

 
$
4.84

 
5.42
 
$
—
Fair Value Assumptions for Stock Options
The fair value was determined by the Black-Scholes option pricing model using the following weighted-average assumptions:
 
Three Months Ended
March 31,
 
2020
 
2019
Expected term, in years
5.90

 
6.36

Expected volatility
89.7
%
 
98.5
%
Risk-free interest rate
0.9
%
 
2.6
%
Expected dividend yield
—
%
 
—
%
Weighted-average exercise price
$
0.89

 
$
2.95
Allocation of Share-based Compensation Expense
Share-based compensation related to all equity awards issued pursuant to the 2008 Plan and 2016 Plan and for estimated shares to be issued under the ESPP for the purchase periods active during each respective period is included in the condensed consolidated statements of operations and comprehensive loss as follows:
 
Three Months Ended
March 31,
 
2020
 
2019
 
(in thousands)
Research and development
$
187

 
$
395

General and administrative
477

 
623

Total share-based compensation expense
$
664

 
$
1,018