Summary of Significant Accounting Policies - Revenue Recognition - Collaboration Revenue (Details) - USD ($)			3 Months Ended		9 Months Ended
	Sep. 24, 2018	Nov. 01, 2017	Sep. 30, 2019	Sep. 30, 2018	Sep. 30, 2019	Sep. 30, 2018
Collaboration Revenue
Written notice period (in days)		90 days
Revenues			$ 972,779	$ 1,132,067	$ 4,268,206	$ 2,820,269
Elanco
Collaboration Revenue
Collaboration revenue			0	0	0	177,389
Knight
Collaboration Revenue
License agreement term (in years)	15 years
Transfer price receivable upon achievement of certain regulatory and sales milestones	$ 18,000,000
Revenues			$ 0	$ 0	$ 0	$ 0