X0306 4 2018-02-20 false 0001583648 PIERIS PHARMACEUTICALS, INC. PIRS 0001714023 Reine Allan C/O PIERIS PHARMACEUTICALS, INC. 255 STATE STREET, 9TH FLOOR BOSTON MA 02109 false true false false Chief Financial Officer Stock Options (Right to buy) 8.56 2018-02-20 4 A false 53833.00 0.00 A 2028-02-20 Common Stock 53833.00 53833.00 D Stock Options (Right to buy) 5.00 2018-02-20 4 A false 50000.00 0.00 A 2027-08-08 Common Stock 50000.00 50000.00 D The option vests as to 25% of the option shares on January 1, 2019 and vests as to an additional 6.25% of the option shares at the end of each successive three-month period thereafter. On August 9, 2017, the reporting person was granted an option to purchase up to a maximum of 50,000 shares of common stock of which 50,000 shares were earned based on the reporting person's achievement of his 2017 personal goals as certified by the Compensation Committee of the Board of Directors on February 20, 2018. The earned shares vest as to 25% on February 20, 2019 and the remainder vests as to an additional 6.25% quarterly thereafter beginning on the last day of the next calendar quarter. /s/ Marc D. Mantell, Attorney-in-fact 2018-02-26