Collaboration Agreements (Tables)

9 Months Ended

Sep. 30, 2015

Organization Consolidation And Presentation Of Financial Statements [Abstract]

Schedule of Revenues from Collaborations

The following table is a summary of the revenue recognized from the Company’s collaborations for the three and nine months ended September 30, 2015 and 2014:

	Three Months Ended September 30,				Nine Months Ended September 30,
	2015		2014		2015		2014
uniQure:
Milestone payment	$	—	$	14	$	—	$	14
Teva:
Recognition of upfront payment		2,940		3,132		8,724		9,252
Research funding		11		84		101		251
Genentech:
Recognition of upfront payment		182		981		542		2,736
Research funding		910		994		2,730		3,249
Milestone payment		250		7,987		250		7,987
Total collaboration revenue	$	4,293	$	13,192	$	12,347	$	23,489