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Stock-based compensation (Tables)
 9 Months Ended
Sep. 30, 2017
Disclosure of Compensation Related Costs, Share-based Payments [Abstract]  
Schedule of Employee Service Share-based Compensation Allocation
Total share-based compensation expense by statement of operations is presented below (in thousands):
 
Three Months Ended September 30,
 
Nine Months Ended September 30,
 
2017
 
2016
 
2017
 
2016
Research and development expense
$
584

 
$
1,361

 
$
2,504

 
$
4,297

General and administrative expense
895

 
1,399

 
2,964

 
3,789

 
$
1,479

 
$
2,760

 
$
5,468

 
$
8,086