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Stockholders' Equity
 6 Months Ended
Jun. 30, 2016
Disclosure of Compensation Related Costs, Share-based Payments [Abstract]  
Stockholders' Equity
Warrants 

As of June 30, 2016 the following warrants for common stock were issued and outstanding:
Issue date
 
Expiration date
 
Exercise price
 
Number of warrants outstanding
November 21, 2012
 
November 21, 2017
 
$
7.86

 
695,383



During the three months ended June 30, 2016, no warrants were exercised. During the six months ended June 30, 2016, warrants for 419,244 shares of the Company's common stock were exercised via cashless exercises and 313,756 shares were exercised for cash generating proceeds of $2.1 million for the six months ended June 30, 2016. The Company issued a total of 603,545 shares of common stock for the same period. During the three and six months ended June 30, 2015, warrants for 179,296 and 192,570 shares of the Company's common stock were exercised via cashless exercises and the Company issued a total of 139,503 and 148,818 shares of common stock, respectively.
Stockholders' Equity

Share-based Compensation

Total share-based compensation expense by operating statement classification is presented below (in thousands):
 
Three Months Ended June 30,
 
Six Months Ended June 30,
 
2016
 
2015
 
2016
 
2015
Research and development expense
$
1,478

 
$
931

 
$
2,936

 
$
1,739

General and administrative expense
810

 
1,861

 
2,390

 
3,369

 
$
2,288

 
$
2,792

 
$
5,326

 
$
5,108


    
During the three months ended June 30, 2016 and 2015, 1,836 and 4,750 shares were issued pursuant to stock option exercises, respectively, generating immaterial net proceeds. During the six months ended June 30, 2016 and 2015, 22,132 and 26,916 shares were issued pursuant to stock option exercises generating net proceeds of $0.2 million and $0.3 million, respectively.