X0306 4 2019-08-30 0 0001512499 LINDBLAD EXPEDITIONS HOLDINGS, INC. LIND 0001574445 Dryden L. Dyson 96 MORTON STREET NEW YORK NY 10014 1 0 0 0 Common Stock 2019-08-30 4 S 0 50251 18.7692 D 1522114 D Common Stock 2019-09-03 4 S 0 92032 18.5965 D 1430082 D Common Stock 2019-09-04 4 S 0 21981 18.7116 D 1408101 D The reported transaction was effected pursuant to a Rule 10b5-1 trading plan adopted by the reporting person. The price in Column 4 is a weighted average price. The prices actually received ranged from $18.48 to $19.13. The reporting person has provided to the issuer, and will provide to any security holder of the issuer, or the SEC staff, upon request, information regarding the number of shares sold at each price within the range for all transactions reported in this Form 4 utilizing a weighted average price. The price in Column 4 is a weighted average price. The prices actually received ranged from $18.5446 to $18.7056. The reporting person has provided to the issuer, and will provide to any security holder of the issuer, or the SEC staff, upon request, information regarding the number of shares sold at each price within the range for all transactions reported in this Form 4 utilizing a weighted average price. /s/ L. Dyson Dryden 2019-09-04