Condensed Consolidated Statements of Operations - USD ($) shares in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2020	Sep. 30, 2019	Sep. 30, 2020	Sep. 30, 2019
Total revenue	$ 157,000	$ 205,000	$ 493,000	$ 558,000
Operating expenses:
Research and development	21,605,000	12,486,000	54,124,000	35,365,000
General and administrative	9,398,000	8,691,000	28,795,000	27,677,000
Restructuring credits	0	0	0	(61,000)
Total operating expenses	31,003,000	21,177,000	82,919,000	62,981,000
Loss from operations	(30,846,000)	(20,972,000)	(82,426,000)	(62,423,000)
Other income (expense):
Interest income	43,000	2,034,000	1,359,000	6,634,000
Other income (expense), net	11,000	(42,000)	3,000	176,000
Total other income, net	54,000	1,992,000	1,362,000	6,810,000
Loss before income taxes	(30,792,000)	(18,980,000)	(81,064,000)	(55,613,000)
Provision for (benefit from) income taxes	(215,000)	468,000	(636,000)	510,000
Net loss	$ (30,577,000)	$ (19,448,000)	$ (80,428,000)	$ (56,123,000)
Basic and diluted net loss per share (in dollars per share)	$ (0.77)	$ (0.49)	$ (2.02)	$ (1.41)
Shares used to compute basic and diluted net loss per share (in shares)	39,917	39,897	39,912	39,877
Collaboration
Total revenue	$ 157,000	$ 205,000	$ 443,000	$ 558,000
License
Total revenue	$ 0	$ 0	$ 50,000	$ 0