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License, Collaboration and Other Significant Agreements - Revenue Recognized Resulting from Changes in Contract Assets and Liabilities (Details) - USD ($)
$ in Thousands
3 Months Ended
Mar. 31, 2022
Mar. 31, 2021
Revenue Recognized in the Period from:    
Amounts included in deferred revenue at the beginning of the period $ 6,602 $ 5,382
Performance obligations satisfied in previous periods $ 0 $ 0