X0306 4 2020-08-18 0 0001497253 ORGANOVO HOLDINGS, INC. ONVO 0001197086 CROUCH TAYLOR 440 STEVENS AVENUE, SUITE 200 SOLANA BEACH CA 92075 1 1 0 0 CEO & President Common Stock 2020-08-18 4 F 0 5510 0.5692 D 125144 D Represents the mandatory sale of shares to satisfy the Reporting Person's tax withholding obligations, in connection with the partial vesting and settlement of restricted stock units granted May 2018, pursuant to the Reporting Person's agreement with the Issuer. /s/ Jennifer Bush, as attorney-in-fact 2020-08-18