X0306 4 2019-11-18 0 0001497253 ORGANOVO HOLDINGS, INC. ONVO 0001197086 CROUCH TAYLOR 6275 NANCY RIDGE DRIVE, SUITE 110 SAN DIEGO CA 92121 1 1 0 0 CEO & President Common Stock 2019-11-18 4 F 0 5075 0.4493 D 143039 D Represents the mandatory sale of shares to satisfy the Reporting Person's tax withholding obligations, in connection with the partial vesting and settlement of restricted stock units granted May 2018, pursuant to the Reporting Person's agreement with the Issuer. /s/ Jennifer Bush, as attorney-in-fact 2019-11-19