X0306 4 2019-08-16 0 0001497253 ORGANOVO HOLDINGS, INC. ONVO 0001621152 Bush Jennifer 6275 NANCY RIDGE DRIVE, SUITE 110 SAN DIEGO CA 92121 0 1 0 0 General Counsel & Secretary Common Stock 2019-08-16 4 F 0 11960 0.2856 D 243833 D Represents the mandatory sale of shares to satisfy the Reporting Person's tax withholding obligations, in connection with the partial vesting and settlement of restricted stock units granted in July 2016, June 2017 and May 2018, pursuant to the Reporting Person's agreement with the Issuer. /s/ Jennifer Bush 2019-08-19