X0306 4 2019-05-16 0 0001497253 ORGANOVO HOLDINGS, INC. ONVO 0001651405 Gallant Paul 6275 NANCY RIDGE DRIVE, SUITE 110 SAN DIEGO CA 92121 0 1 0 0 General Manager Common Stock 2019-05-16 4 F 0 9274 0.97 D 200835 D Represents the mandatory sale of shares to satisfy the Reporting Person's tax withholding obligations, in connection with the partial vesting and settlement of restricted stock units granted in July 2016, June 2017 and May 2018, pursuant to the Reporting Person's agreement with the Issuer. This number does not include 1,036,667 shares of common stock underlying outstanding options held by Mr. Gallant; post-transaction Mr. Gallant beneficially owns or has a right to acquire 1,237,502 shares of common stock. /s/ Jennifer Bush, as attorney-in-fact 2019-05-20