X0306 4 2018-02-16 0 0001497253 ORGANOVO HOLDINGS, INC. ONVO 0001197083 KUSSMAN CRAIG 6275 NANCY RIDGE DRIVE, SUITE 110 SAN DIEGO CA 92121 0 1 0 0 Chief Financial Officer Common Stock 2018-02-16 4 F 0 13265 1.0855 D 300483 D Represents the mandatory sale of shares to satisfy the Reporting Person's tax withholding obligations, in connection with the partial vesting and settlement of restricted stock units granted in August 2016 and June 2017, pursuant to the Reporting Person's agreement with the Issuer. This number does not include 330,000 shares of common stock underlying oustanding options held by Mr. Kussman; post-transaction Mr. Kussman continues to beneficially own or have a right to acquire 630,483 shares of common stock. /s/ Craig Kussman 2018-02-20