X0306 4 2016-11-16 0 0001497253 ORGANOVO HOLDINGS, INC. ONVO 0001549671 David Eric 6275 NANCY RIDGE DRIVE, SUITE 110 SAN DIEGO CA 92121 0 1 0 0 CSO & EVP Pre-Clinical Dev Common Stock 2016-11-16 4 F 0 1745 3.2594 D 703356 D Represents the mandatory sale of shares to satisfy the Reporting Person's tax withholding obligations, in connection with the partial vesting and settlement of restricted stock units granted in July 2016, pursuant to the Reporting Person's agreement with the Issuer. This number does not include 1,070,000 shares of common stock underlying outstanding options held by Dr. David; post-transaction Dr. David continues to beneficially own or have a right to acquire 1,773,356 shares of common stock. /s/ Eric David 2016-11-18