X0306 4 2016-11-16 0 0001497253 ORGANOVO HOLDINGS, INC. ONVO 0001621152 Bush Jennifer 6275 NANCY RIDGE DRIVE, SUITE 110 SAN DIEGO CA 92121 0 1 0 0 General Counsel & Secretary Common Stock 2016-11-16 4 F 0 1210 3.2594 D 47576 D Represents the mandatory sale of shares to satisfy the Reporting Person's tax withholding obligations, in connection with the partial vesting and settlement of restricted stock units granted in July 2016, pursuant to the Reporting Person's agreement with the Issuer. This number does not include 375,000 shares of common stock underlying outstanding options held by Ms. Bush; post-transaction Ms. Bush continues to beneficially own or have a right to acquire 422,576 shares of common stock. /s/ Jennifer Bush 2016-11-18