X0306 4 2016-08-17 0 0001497253 ORGANOVO HOLDINGS, INC. ONVO 0001364590 Maroun Richard E 6275 NANCY RIDGE DRIVE SAN DIEGO CA 92121 1 0 0 0 Common Stock 2016-08-17 4 A 0 7500 0.00 A 7500 D Common Stock 2016-08-17 4 A 0 7500 0.00 A 15000 D Stock Option (Right to Buy) 4.35 2016-08-17 4 A 0 18500 0.00 A 2025-08-17 Common Stock 18500 18500 D Stock Option (Right to Buy) 4.35 2016-08-17 4 A 0 18500 0.00 A 2025-08-17 Common Stock 18500 18500 D Initial long-term equity award issued pursuant to the Issuer's non-employee director compensation plan. The shares represent a grant of Restricted Stock Units ("RSUs"). The RSUs represent a right to receive shares of the Issuer's common stock, at no additional conversion or exercise price. The RSUs vest and settle over 12 equal quarterly installments measured from August 15, 2016. Annual long-term equity award issued pursuant to the Issuer's non-employee director compensation plan. The shares represent a grant of Restricted Stock Units ("RSUs"). The RSUs represent a right to receive shares of the Issuer's common stock, at no additional conversion or exercise price. The RSUs vest and settle in full on the earlier of (I) August 15, 2017 or (ii) immediately prior to the Issuer's next annual stockholder meeting. The option shares vest and become exercisable over 12 equal quarterly installments measured from August 15, 2016. The option shares vest and become exercisable in full on the earlier of (i) August 15, 2017 or (ii) immediately prior to the Issuer's next annual stockholder meeting. /s/ Richard Maroun 2016-08-19