Exhibit 107

Calculation of Filing Fee Tables

Form S-1
(Form Type)Organovo Holdings, Inc.
(Exact Name of Registrant as Specified in its Charter)Table 1: Newly Registered and Carry Forward Securities

Security Type

Security Class Title(1)

Fee Calculation or Carry Forward Rule

Amount Registered

Proposed Maximum Offering Price per Unit

Maximum Aggregate Offering Price

Fee Rate

Amount of Registration Fee

Carry Forward Form Type

Carry Forward File Number

Carry Forward Initial Effective Date

Filing Fee Previously Paid in Connection with Unsold Securities to be Carried Forward

Newly Registered Securities

Fees to Be Paid

Equity

Common Stock, par value $0.001 per share or Pre-Funded warrants to purchase shares of Common Stock

Rule 457(o)

___

___

$10,000,000(2)

0.00014760

$1,476.00

Equity

Common Stock underlying the Pre-Funded Warrants

Rule 457(o)

___

___

Included above

___

Equity

Common warrants to purchase shares of Common Stock

Rule 457(o)

Included below

Equity

Common Stock underlying the common warrants(3)

Rule 457(g)

$10,000,000

0.00014760

$1,476.00

Fees Previously Paid

Equity

Common Stock, par value $0.001 per share or Pre-Funded warrants to purchase shares of Common Stock

Rule 457(o)

$10,000,000

0.00014760

$1,476.00

Equity

Common Stock underlying the Pre-Funded Warrants

Rule 457(o)

___

Included above

___

Carry Forward Securities

Carry Forward Securities

N/A

___

N/A

N/A

N/A

N/A

Total Offering Amounts

$11,500,000

$2,952.00

Total Fees Previously Paid

0.00014760

$1,476.00

Total Fee Offsets

Net Fee Due

$1,476.00

(1) The Registrant may issue pre-funded warrants to purchase common stock in the offering. The purchase price of each pre-funded warrant will equal the price per share at which common stock are being sold to the public in this offering, minus $0.001, which constitutes the pre-funded portion of the exercise price, and the remaining unpaid exercise price of the pre-funded warrant will equal $0.001 per share (subject to adjustment as provided for therein). The proposed maximum aggregate offering price of the common stock proposed to be sold in the offering will be reduced on a dollar-for-dollar basis based on the offering price of any pre-funded warrants offered and sold in the offering, and the proposed maximum offering price of the pre-funded warrants to be sold in the offering will be reduced on a dollar-for-dollar basis based on the offering price of any common stock sold in the offering.

(2) Estimated solely for the purpose of calculating the amount of the registration fee pursuant to Rule 457(o) under the Securities Act of 1933, as amended (the “Securities Act”). Pursuant to Rule 416 under the Securities Act, this registration statement also covers any additional securities that may

be offered or issued in connection with any stock split, stock dividend or pursuant to anti-dilution provisions of any of the securities. The proposed maximum aggregate offering price of the common stock, pre-funded warrants and common stock issuable upon exercise of the pre-funded warrants, if any, is $10,000,000.

(3) In accordance with Rule 457(g) under the Securities Act, because the shares of the common stock underlying the common warrants are registered hereby, no separate registration fee is required with respect to such common warrants..