Subsequent Events - Additional Information (Details) - Subsequent Event - Sato License Agreement	Jan. 19, 2017 USD ($)	Jan. 12, 2017 USD ($)	Jan. 12, 2017 JPY (¥)
Subsequent Event [Line Items]
Upfront payment under license agreement | ¥			¥ 1,250,000,000
License agreement additional term		2 years
Description of license terms		The term of the Agreement and the period during which Sato must pay royalties under the Agreement expires, on a licensed product-by-licensed product basis, on the tenth anniversary of the first commercial sale of a licensed product in the licensed field in the licensed territory.
Maximum potential amount of research activity costs to be paid as per the agreement | $		$ 1,000,000
Proceed from license agreement | $	$ 10,813,000
License agreement execution expense paid to third party | $		$ 216,000
Maximum
Subsequent Event [Line Items]
Aggregate Development And Regulatory Milestone Payments Potentially Receivable Under License Agreement | ¥			2,750,000,000
Aggregate Commercial Milestone Payments Potentially Receivable Under License Agreement | ¥			¥ 900,000,000