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Collaboration Agreements - Other Revenue (Details) - USD ($)
3 Months Ended
Mar. 31, 2022
Mar. 31, 2021
Collaboration agreements    
Collaboration revenue $ 2,036,000 $ 11,000
Asana BioSciences    
Collaboration agreements    
Next potential milestone payment eligible to receive 2,500,000  
Asana BioSciences | Limited services    
Collaboration agreements    
Collaboration revenue $ 300,000 $ 0