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Collaboration and License Agreements - Schedule of Revenues as a Result of Changes in Contract Liabilities (Details) - USD ($)
$ in Thousands
3 Months Ended 9 Months Ended
Sep. 30, 2019
Sep. 30, 2018
Sep. 30, 2019
Sep. 30, 2018
Revenue Recognition and Deferred Revenue [Abstract]        
Amounts included in the contract liability at the beginning of the period $ 4,404 $ 7,555 $ 28,823 $ 23,472
Performance obligations satisfied in previous periods $ 0 $ 219 $ 0 $ 762