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Collaboration and Licensing Agreements - Deferred Revenue (Details) - USD ($)
$ in Thousands
3 Months Ended
Mar. 31, 2023
Mar. 31, 2022
Deferred revenue - current    
Beginning balance $ 4,211  
Ending balance 5,545  
Deferred revenue - non-current    
Beginning balance 9,025  
Ending balance 11,498  
2017 KKC Agreement    
Deferred revenue - current    
Beginning balance 4,211 $ 0
Increases due to amounts reclassified from non-current, to be recognized in the next twelve months 809
Increases to amounts invoiced, for which cash has not yet been received 525 0
Ending balance 5,545 0
Deferred revenue - non-current    
Beginning balance 9,025 4,727
Increases to amounts invoiced, for which cash has not yet been received 3,282 3,829
Increase due to unbilled prepayments recorded during the period 0 7
Decreases due to amounts reclassified as current, to be recognized in the next twelve months (809) 0
Ending balance $ 11,498 $ 8,563