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Selected Quarterly Financial Information (unaudited)
 12 Months Ended
Dec. 31, 2016
Quarterly Financial Information Disclosure [Abstract]  
Selected Quarterly Financial Information (unaudited)
14. Selected Quarterly Financial Information (unaudited)
The following is a summary of the quarterly results of operations for the years ended December 31, 2016 and 2015:
 
2016
 
March 31
 
June 30
 
September 30
 
December 31
Collaboration revenue
$
15,721

 
$
28,198

 
$
1,668

 
$
5,322

Research and development expenses
37,770

 
48,262

 
50,854

 
59,409

General and administrative expenses
14,696

 
10,489

 
12,024

 
12,968

Loss from operations
(36,745
)
 
(30,553
)
 
(61,210
)
 
(67,055
)
Net loss attributable to common stockholders
$
(36,301
)
 
$
(29,945
)
 
$
(60,891
)
 
$
(66,283
)
Basic and diluted loss per common share
$
(1.03
)
 
$
(0.85
)
 
$
(1.71
)
 
$
(1.86
)

 
2015
 
March 31
 
June 30
 
September 30
 
December 31
Collaboration revenue
$
41,678

 
$
1,597

 
$
3,448

 
$
4,782

Research and development expenses
24,557

 
32,059

 
40,479

 
33,917

General and administrative expenses
9,584

 
11,959

 
10,412

 
12,066

Income (loss) from operations
7,537

 
(42,421
)
 
(47,443
)
 
(41,201
)
Net income (loss) attributable to common stockholders
$
6,636

 
$
(37,131
)
 
$
(39,573
)
 
$
(35,649
)
Basic earnings (loss) per common share
$
0.19

 
$
(1.08
)
 
$
(1.14
)
 
$
(1.02
)
Diluted earnings (loss) per common share
$
0.19

 
$
(1.08
)
 
$
(1.14
)
 
$
(1.02
)