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Share-Based Compensation - (Tables)
 3 Months Ended
Mar. 31, 2021
Share-based Payment Arrangement [Abstract]  
Schedule of Allocation of Share-based Compensation Costs by Award Type Stock-based compensation by award type was as follows:
 Three Months Ended March 31,
 20212020
Stock options$1,469 $1,374 
Restricted stock units792 936 
Employee stock purchase plan31 14 
Total$2,292 $2,324 
Schedule of Allocation of Share-based Compensation Costs by Financial Statement Line
The Company allocated stock-based compensation expense in the Company’s Consolidated Statements of Operations and Comprehensive Loss as follows:
 Three Months Ended March 31,
 20212020
Research and development$1,343 $1,153 
General and administrative949 1,171 
Total$2,292 $2,324 
Summary of the stock option activity including weighted average exercise prices, options outstanding and exercisable
A summary of the stock option activity, weighted average exercise prices, options outstanding, exercisable and expected to vest as of March 31, 2021 is as follows (in thousands except weighted average exercise price):
 Number of Shares Underlying OptionsWeighted
Average
Exercise
Price
Outstanding, December 31, 20208,928 $9.22 
Granted266 $6.38 
Exercised(2)$3.80 
Forfeited(14)$4.77 
Outstanding, March 31, 20219,178 $9.41 
Vested and exercisable, March 31, 20214,856 $13.22 
Vested and expected to vest, March 31, 20218,832 $9.58 
Summary of the Company's outstanding shares of RSU awards
The following table presents a summary of the Company's outstanding RSU awards granted as of March 31, 2021 (in thousands except weighted average grant-date fair value):
Restricted
Stock
Units		Weighted Average
Grant-Date
Fair Value
Outstanding, December 31, 20201,958 $6.95 
Awarded112 $6.33 
Vested(22)$3.83 
Forfeited(10)$7.50 
Outstanding, March 31, 20212,038 $7.01 
Outstanding, Expected to vest1,815 $5.31