Restructuring Activities (Tables)

6 Months Ended

Jun. 30, 2018

Restructuring and Related Activities [Abstract]

Schedule of severance costs

The following is a reconciliation of the severance-related accrual activity for the six months ended June 30, 2018:

	Accrued Severance and Other Employee Costs
Beginning Balance	$	2,529
Accrued restructuring expenses	—
Payments	(2,082		)
Ending Balance	$	447

In connection with the Company's restructuring activities, the Company recognized severance, stock compensation, other employee costs and lease termination costs,all of which have been reported in the Company’s Statements of Operations, as follows:

	Three Months Ended June 30,				Six Months Ended June 30,
	2018		2017		2018		2017
Research and development	$	—	$	1,048	$	—	$	5,883
General and administrative	—		708		—		4,649
Total	$	—	$	1,756	$	—	$	10,532