X0306 4 2017-04-03 0 0001098972 AGENUS INC agen 0001383313 Wright Timothy C/O AGENUS INC 3 FORBES ROAD LEXINGTON MA 02421 1 0 0 0 Deferred Shares 4.037 2017-04-03 4 A 0 1586.909 4.037 A Common Stock 1586.909 1586.909 D Acquired under the Agenus Inc. Director Deferred Compensation Plan, as amended. Deferred Shares represent an equal amount of the Company's common stock to be distributed under the terms of the Plan, typically once the director ceases to serve as a director of the Company. Christine M. Klaskin, by Power of Attorney 2017-04-06