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Stock-Based Compensation
3 Months Ended
Mar. 31, 2022
Share-based Payment Arrangement [Abstract]  
Stock-Based Compensation Stock-Based CompensationStock-based compensation expense for stock options and RSUs is reflected in the condensed consolidated statements of operations and comprehensive loss as follows (in thousands):
Three months ended March 31,
20222021
General and administrative$2,216 $590 
Research and development1,083 243 
Total$3,299 $833 
As of March 31, 2022, the Company had $27.9 million of unrecognized stock-based compensation expense related to stock options and RSUs outstanding. This expense is expected to be recognized over a weighted-average period of 2.3 years as of March 31, 2022.
Equity Plans
The Company maintains two equity compensation plans, the 2014 Ocugen OpCo, Inc. Stock Option Plan (the "2014 Plan") and the Ocugen, Inc. 2019 Equity Incentive Plan (the "2019 Plan", collectively with the 2014 Plan, the "Plans"). As of March 31, 2022, the 2014 Plan and 2019 Plan authorize for the granting of up to 0.8 million and 19.5 million equity awards with respect to the Company's common stock, respectively. In addition to stock options and RSUs granted under the Plans, the Company has granted certain stock options and RSUs as material inducements to employment in accordance with Nasdaq Listing Rule 5635(c)(4), which were granted outside of the Plans.
Options to Purchase Common Stock
The following table summarizes the stock option activity:
Number of SharesWeighted-Average Exercise PriceWeighted Average Remaining Contractual LifeAggregate Intrinsic Value
Options outstanding at December 31, 2021
10,086,167 $2.59 8.8$24,664 
Granted4,323,937 $4.42 
Exercised(277,323)$0.65 
Forfeited(130,327)$6.48 
Options outstanding at March 31, 2022
14,002,454 $3.16 8.9$14,468 
Options exercisable at March 31, 2022
2,913,960 $2.37 8.3$4,431 
There were 1.2 million of stock options with performance-based vesting conditions outstanding as of March 31, 2022 and December 31, 2021, of which 0.9 million were not yet vested and exercisable as of March 31, 2022 and December 31, 2021. The weighted average grant date fair values of stock options granted during the three months ended March 31, 2022 and 2021 were $3.61 and $1.73, respectively. The total fair values of stock options vested during the three months ended March 31, 2022 and 2021 were $2.8 million and $0.3 million, respectively.
RSUs
The following table summarizes the RSU activity:
Number of SharesWeighted-Average Grant-Date Fair Value
RSUs outstanding at December 31, 2021
191,811 $6.79 
Granted1,130,270 $4.45 
Forfeited(20,812)$6.36 
RSUs outstanding at March 31, 2022
1,301,269 $4.76