Income Taxes - Schedule of Effective Income Tax Rate Reconciliation (Details) - USD ($) $ in Thousands	12 Months Ended
	Dec. 31, 2019	Dec. 31, 2018	Dec. 31, 2017
Income Tax Disclosure [Abstract]
Computed statutory income tax benefit from continuing operations	$ (43,824)	$ (90,246)	$ (26,645)
State and provincial income tax benefit, net of federal income taxes	(7,298)	(23,347)	(1,888)
Nondeductible stock based compensation	10,303	4,696	3,391
Nondeductible officer compensation	595	294	476
Gain on dividend distribution of AquaBounty common stock	0	0	3,965
Impairment of goodwill	273	0	4,700
Research and development tax incentives	(1,772)	(185)	(359)
Acquisition and internal restructuring transaction costs	260	52	354
Provisional impact of the Tax Act	0	0	85,288
Enacted changes in foreign tax rates and foreign tax reforms	0	0	2,021
Reacquired in-process research and development	0	2,696	0
Change in deferred state tax rate	0	8,666	0
United States-foreign rate differential	(76)	215	410
Other, net	(72)	(3,517)	(189)
Income tax reconciliation income tax benefit before valuation allowance, total	(41,611)	(100,676)	71,524
Change in valuation allowance for deferred tax assets	40,681	85,251	(69,452)
Total income tax expense (benefit) from continuing operations	$ (930)	$ (15,425)	$ 2,072