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Property, Plant and Equipment, Net - Schedule of Property, Plant and Equipment (Details) - USD ($)
$ in Thousands
Dec. 31, 2019
Dec. 31, 2018
Property, Plant and Equipment [Abstract]    
Land and land improvements $ 9,814 $ 10,001
Buildings and building improvements 11,765 20,099
Furniture and fixtures 1,315 1,812
Equipment 54,448 63,947
Leasehold improvements 12,821 14,219
Breeding stock 5,191 4,582
Computer hardware and software 9,434 10,789
Construction and other assets in progress 5,313 10,497
Property, plant and equipment, gross 110,101 135,946
Less: Accumulated depreciation and amortization (49,132) (49,050)
Property, plant and equipment, net $ 60,969 $ 86,896