Collaboration and Licensing Revenue - Summary of Deferred Revenue (Details) - USD ($) $ in Thousands	Dec. 31, 2019	Dec. 31, 2018
Deferred Revenue Arrangement [Line Items]
Deferred revenue	$ 53,833	$ 57,816
Current portion of deferred revenue	5,697	11,088
Long-term portion of deferred revenue	48,136	46,728
Collaboration and licensing agreements
Deferred Revenue Arrangement [Line Items]
Deferred revenue	50,593	54,323
Prepaid product and service revenues
Deferred Revenue Arrangement [Line Items]
Deferred revenue	2,805	2,933
Other
Deferred Revenue Arrangement [Line Items]
Deferred revenue	$ 435	$ 560