Investments in Joint Ventures - Intrexon T1D Partners - Additional Information (Details)	1 Months Ended		12 Months Ended
	Nov. 30, 2018 USD ($) shares	Mar. 31, 2016 USD ($)	Dec. 31, 2019 USD ($)	Dec. 31, 2018 USD ($)	Dec. 31, 2017 USD ($)	Oct. 31, 2018 board_seat
Schedule Of Investments In Joint Venture [Line Items]
Shares issued for reacquired in-process research and development				$ 159,323,000
Deferred revenue			$ 53,833,000	57,816,000
Reacquisition of in-process research and development			0	236,748,000	$ 0
Collaboration and licensing agreements
Schedule Of Investments In Joint Venture [Line Items]
Deferred revenue			$ 50,593,000	$ 54,323,000
Intrexon T1D Partners, LLC
Schedule Of Investments In Joint Venture [Line Items]
Membership interest	100.00%	50.00%
Maximum additional capital contributions committed		$ 5,000,000
Total number of seats on the joint venture's governing board, internally selected | board_seat						2
Total number of seats on the joint venture's governing board, externally selected | board_seat						3
Shares issued in asset acquisition or for reacquired in-process research and development, shares | shares	1,933,737
Shares issued for reacquired in-process research and development	$ 18,970,000
Deferred revenue	8,517,000
Reacquisition of in-process research and development	$ 10,453,000
Investor | Intrexon T1D Partners, LLC
Schedule Of Investments In Joint Venture [Line Items]
Membership interest		50.00%
Capital contribution		$ 10,000,000
Maximum additional capital contributions committed		5,000,000
Intrexon T1D Partners, LLC | Collaboration and licensing agreements
Schedule Of Investments In Joint Venture [Line Items]
Collaborative agreement, consideration received, value		$ 10,000,000