Mergers and Acquisitions - Asset Acquisition of Certain Harvest Entities (Details) - USD ($)	1 Months Ended	12 Months Ended
	Sep. 30, 2018	Dec. 31, 2019	Dec. 31, 2018	Dec. 31, 2017	Aug. 31, 2018
Business Acquisition [Line Items]
Long-term debt issued to a related party in an asset acquisition		$ 0	$ 30,000,000	$ 0
Cash received in asset acquisition		0	15,500,000	(14,219,000)
Investments in affiliates		1,461,000	2,139,000
Deferred revenue		53,833,000	57,816,000
Reacquisition of in-process research and development		$ 0	$ 236,748,000	$ 0
Harvest Intrexon Enterprise Fund I, LP
Business Acquisition [Line Items]
Cash received in asset acquisition	$ 15,500,000
Precigen ActoBio, Inc. | Harvest Intrexon Enterprise Fund I, LP
Business Acquisition [Line Items]
Long-term debt issued to a related party in an asset acquisition	30,000,000
CRS Bio, Inc.; Genten Therapeutics, Inc.; and Relive Genetics, Inc.
Business Acquisition [Line Items]
Investments in affiliates					$ 4,303,000
Ownership interest		100.00%
Deferred revenue	10,078,000
Reacquisition of in-process research and development	$ 8,721,000