Property, Plant and Equipment, Net

12 Months Ended

Dec. 31, 2019

Property, Plant and Equipment [Abstract]

Property, Plant and Equipment, Net

Property, Plant and Equipment, Net

Property, plant and equipment consist of the following:

	December 31,
	2019			2018
Land and land improvements	$	9,814		$	10,001
Buildings and building improvements	11,765			20,099
Furniture and fixtures	1,315			1,812
Equipment	54,448			63,947
Leasehold improvements	12,821			14,219
Breeding stock	5,191			4,582
Computer hardware and software	9,434			10,789
Construction and other assets in progress	5,313			10,497
	110,101			135,946
Less: Accumulated depreciation and amortization	(49,132		)	(49,050		)
Property, plant and equipment, net	$	60,969		$	86,896

The deconsolidation of AquaBounty (Note 1) in April 2019 resulted in the reduction of $24,186 of property, plant and equipment, net on the accompanying consolidated balance sheet during the year ended December 31, 2019.

During the year ended December 31, 2019, the Company recorded $448 of property, plant and equipment impairment losses in conjunction with the closing of two of its operating units during the third quarter of 2019.

Depreciation expense was $11,869, $12,439 and $10,137 for the years ended December 31, 2019, 2018 and 2017, respectively.