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Related Party Transactions - Additional Information (Details) - USD ($)
$ in Thousands
1 Months Ended 12 Months Ended
Nov. 30, 2021
Dec. 31, 2020
Dec. 31, 2022
Dec. 31, 2021
Dec. 31, 2020
Nov. 30, 2020
Related Party Transaction [Line Items]            
Shares issued in conjunction with settlement agreement         $ 18,103  
Deferred revenue     $ 1,843 $ 24,513    
Exotech, Inc.; AD Skincare, Inc.; and Thrive Agrobiotics, Inc.            
Related Party Transaction [Line Items]            
Investments in affiliates           $ 326
Collaboration and licensing agreements            
Related Party Transaction [Line Items]            
Deferred revenue     1,818 23,023    
Harvest Intrexon Enterprise Fund I, LP            
Related Party Transaction [Line Items]            
Stock issued in conjunction with settlement agreement, shares   2,117,264        
Shares issued in conjunction with settlement agreement   $ 18,103        
Loss related to litigation settlement   11,436        
Exotech, Inc.; AD Skincare, Inc.; and Thrive Agrobiotics, Inc. | Collaboration and licensing agreements            
Related Party Transaction [Line Items]            
Deferred revenue   $ 6,993     6,993  
Third Security            
Related Party Transaction [Line Items]            
Expense for services     25 100 159  
Sublease rental income     $ 0 $ 75 $ 83  
Cash received on contract termination $ 143          
Third Security | Executive Chairman            
Related Party Transaction [Line Items]            
Ownership interest     100.00%