Lines of Credit and Long-Term Debt - Long-Term Debt - Additional Information (Details)	1 Months Ended	3 Months Ended
	Jul. 31, 2018 USD ($) day $ / shares	Mar. 31, 2022 USD ($)	Mar. 31, 2021 USD ($)	Jan. 01, 2022 USD ($)	Dec. 31, 2021 USD ($)
Debt Instrument [Line Items]
Deferred income taxes		$ (58,000)	$ (56,000)
Long-term debt, net of current portion		201,112,000			$ 182,749,000
Accrued interest		1,750,000
Long-term debt		201,467,000			183,151,000
Cumulative Effect, Period of Adoption, Adjustment | Accounting Standards Update 2020-06 Retrospective
Debt Instrument [Line Items]
Long-term debt, net of current portion				$ 18,196,000
Cumulative Effect, Period of Adoption, Adjustment | Accounting Standards Update 2020-06 Retrospective | Additional Paid-in Capital
Debt Instrument [Line Items]
Stockholders' equity, including portion attributable to noncontrolling interest				36,868,000
Cumulative Effect, Period of Adoption, Adjustment | Accounting Standards Update 2020-06 Retrospective | Accumulated Deficit
Debt Instrument [Line Items]
Stockholders' equity, including portion attributable to noncontrolling interest				$ 18,672,000
Convertible debt
Debt Instrument [Line Items]
Long-term debt		198,362,000			179,882,000
Notes payable
Debt Instrument [Line Items]
Long-term debt		$ 3,105,000			3,217,000
Trans Ova Genetics, LC | Notes payable | American State Bank
Debt Instrument [Line Items]
Debt instrument, interest rate, stated percentage		3.95%
Long-term debt		$ 3,105,000
Debt instrument, periodic payment		39,000
3.5% Convertible Notes Due 2023
Debt Instrument [Line Items]
Aggregate principal amount	$ 200,000,000	$ 200,000,000
Proceeds from long-term debt, net of issuance costs	193,958,000
Debt issuance costs	$ 6,042,000
Debt instrument, interest rate, stated percentage	3.50%	3.50%
Conversion rate	58.6622
Principal amount used in conversion	$ 1,000
Conversion price (in usd per share) | $ / shares	$ 17.05
Debt instrument redemption price, percentage	100.00%
Carrying value of convertible debt	$ 143,723,000	$ 198,362,000			$ 179,882,000
Equity component of convertible debt, net of issuance costs and deferred taxes	50,235,000
Deferred income taxes	$ 13,367,000
Effective interest rate on convertible notes		425.00%
Convertible notes, unamortized discount and issuance costs		$ 1,638,000
3.5% Convertible Notes Due 2023 | Minimum
Debt Instrument [Line Items]
Common stock price trading days | day	20
Common stock price consecutive trading days | day	30
Percentage of common share price over conversion price for conversion	130.00%
3.5% Convertible Notes Due 2023 | Maximum
Debt Instrument [Line Items]
Common stock price trading days | day	5
Common stock price consecutive trading days | day	5
Debt instrument redemption price, percentage	98.00%