Collaboration and Licensing Revenue - Summary of Deferred Revenue (Details) - USD ($) $ in Thousands	Mar. 31, 2022	Dec. 31, 2021
Contract With Customer, Asset And Liability [Line Items]
Long-term portion of deferred revenue	$ 23,023	$ 23,023
Deferred revenue	25,692	27,465
Current portion of deferred revenue	2,669	4,442
Intrexon Energy Partners L.L. and Intrexon Energy Partners II, L.L.C
Contract With Customer, Asset And Liability [Line Items]
Long-term portion of deferred revenue	21,205
Collaboration and licensing agreements
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	23,023	23,023
Prepaid product and service revenues
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	2,511	4,229
Other
Contract With Customer, Asset And Liability [Line Items]
Deferred revenue	$ 158	$ 213