Lines of Credit and Long-Term Debt - Long-Term Debt - Additional Information (Details)	1 Months Ended			12 Months Ended
	Oct. 31, 2020 shares	Sep. 30, 2020 shares	Jul. 31, 2018 USD ($) day $ / shares	Dec. 31, 2021 USD ($)	Dec. 31, 2020 USD ($)	Dec. 31, 2019 USD ($)	Dec. 31, 2018 USD ($)	Sep. 30, 2018 USD ($)
Debt Instrument [Line Items]
Proceeds from long-term debt, net of issuance costs				$ 0	$ 0	$ 376,000
Deferred income taxes				(167,000)	(156,000)	(3,674,000)
Interest payable				3,500,000
Long-term debt				183,151,000	171,882,000
Convertible debt
Debt Instrument [Line Items]
Long-term debt				$ 179,882,000	168,147,000
Convertible debt | Precigen ActoBio, Inc. | Harvest Intrexon Enterprise Fund I, LP
Debt Instrument [Line Items]
Debt instrument, interest rate, stated percentage								3.00%
Interest expense					616,000	$ 921,000
Notes payable to banks | Trans Ova Genetics, LC | American State Bank
Debt Instrument [Line Items]
Debt instrument, interest rate, stated percentage				3.95%
Long-term debt				$ 3,217,000
Debt instrument, periodic payment				39,000
3.5% Convertible Notes Due 2023
Debt Instrument [Line Items]
Aggregate principal amount			$ 200,000,000	200,000,000
Proceeds from long-term debt, net of issuance costs			193,958,000
Debt issuance costs			$ 6,042,000
Debt instrument, interest rate, stated percentage			3.50%
Conversion rate			58.6622
Principal amount used in conversion			$ 1,000
Conversion price (in usd per share) | $ / shares			$ 17.05
Debt instrument redemption price, percentage			100.00%
Carrying value of convertible debt			$ 143,723,000	$ 179,882,000	$ 168,147,000
Adjustments to additional paid in capital, equity component of convertible debt			50,235,000
Deferred income taxes			$ 13,367,000
Effective interest rate on convertible notes				11.02%
Convertible notes, unamortized discount and issuance costs				$ 20,118,000
3.5% Convertible Notes Due 2023 | Minimum
Debt Instrument [Line Items]
Common stock price trading days | day			20
Common stock price consecutive trading days | day			30
Percentage of common share price over conversion price for conversion			130.00%
3.5% Convertible Notes Due 2023 | Maximum
Debt Instrument [Line Items]
Common stock price trading days | day			5
Common stock price consecutive trading days | day			5
Debt instrument redemption price, percentage			98.00%
Merck Convertible Note
Debt Instrument [Line Items]
Aggregate principal amount							$ 25,000,000
Common stock issued upon conversion of long-term debt (in shares) | shares	6,758,400
ActoBio Notes
Debt Instrument [Line Items]
Common stock issued upon conversion of long-term debt (in shares) | shares		6,293,402
Harvest Intrexon Enterprise Fund I, LP | Precigen ActoBio, Inc.
Debt Instrument [Line Items]
Aggregate principal amount								$ 30,000,000