Collaboration and Licensing Revenue (Tables)

12 Months Ended

Dec. 31, 2021

Organization, Consolidation and Presentation of Financial Statements [Abstract]

Summarized Collaboration and Licensing Revenues

The following table summarizes the amounts recorded as revenue in the consolidated statements of operations for each significant counterparty to a collaboration or licensing agreement for the years ended December 31, 2021, 2020, and 2019.

	Year Ended December 31,
	2021		2020		2019
Alaunos Therapeutics, Inc.	$	100	$	200	$	2,171
Oragenics, Inc.	—		3,053		(564)
Intrexon Energy Partners, LLC	—		—		2,596
Intrexon Energy Partners II, LLC	—		—		1,217
Castle Creek Biosciences, Inc.	388		17,810		3,713
Harvest start-up entities (1)	—		—		4,862
Other	18		145		64
Total (2)	$	506	$	21,208	$	14,059

(1)For the year ended December 31, 2019, revenue recognized from collaborations with Harvest start-up entities include Thrive Agrobiotics, Inc.; Exotech Bio, Inc.; and AD Skincare, Inc.

(2)Collaboration and licensing revenues recognized for the years ended December 31, 2021, 2020, and 2019, include the recognition of $397, $20,205, and $7,505, respectively, associated with upfront and milestone payments which were previously deferred.

Summary of Deferred Revenue

Deferred revenue consisted of the following:

	December 31,
	2021		2020
Collaboration and licensing agreements	$	23,023	$	23,420
Prepaid product and service revenues	4,229		2,126
Other	213		277
Total	$	27,465	$	25,823
Current portion of deferred revenue	$	4,442	$	2,800
Long-term portion of deferred revenue	23,023		23,023
Total	$	27,465	$	25,823

Summary of Deferred Revenue by Collaborator

The following table summarizes the remaining balance of deferred revenue associated with upfront and milestone payments for each significant counterparty to a collaboration or licensing agreement as of December 31, 2021 and 2020, as well as the estimated remaining performance period as of December 31, 2021.

	Average Remaining Performance Period (Years)	December 31,
		2021		2020
Intrexon Energy Partners, LLC	2.2	8,362		8,362
Intrexon Energy Partners II, LLC	2.9	12,843		12,843
Castle Creek Biosciences, Inc.	0.0	—		379
Other	2.8	1,818		1,836
Total		$	23,023	$	23,420