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Property, Plant and Equipment, Net
 12 Months Ended
Dec. 31, 2021
Property, Plant and Equipment [Abstract]  
Property, Plant and Equipment, Net Property, Plant and Equipment, Net
Property, plant and equipment consist of the following:
 December 31,
 20212020
Land and land improvements$11,118 $9,844 
Buildings and building improvements12,896 12,088 
Furniture and fixtures1,162 1,228 
Equipment33,584 31,150 
Leasehold improvements4,822 6,260 
Breeding stock1,000 868 
Computer hardware and software5,131 5,684 
Construction and other assets in progress3,824 2,754 
73,537 69,876 
Less: Accumulated depreciation and amortization(39,222)(34,952)
Property, plant and equipment, net$34,315 $34,924 
Depreciation expense was $6,202, $7,449, and $8,222 for the years ended December 31, 2021, 2020, and 2019, respectively.
During the year ended December 31, 2021, the Company recorded impairment losses of $543, which is included in impairment of other noncurrent assets on the accompanying consolidated statement of operations, primarily related to a right-of-use asset at one of the Company's leased locations.
During the year ended December 31, 2020, the Company recorded impairment losses of $920, which is included in impairment of other noncurrent assets on the accompanying consolidated statement of operations, primarily related to right-of-use assets at certain of the Company's leased locations.
During the year ended December 31, 2019, the Company recorded impairment losses of $448, which is included in impairment of other noncurrent assets on the accompanying consolidated statement of operations, related to the impairment of property, plant and equipment in conjunction with the closing of two of its operating units during the third quarter of 2019.