Related Party Transactions - Additional Information (Details) - USD ($) $ in Thousands	1 Months Ended					12 Months Ended
	Nov. 30, 2021	Dec. 31, 2020	Jan. 31, 2020	Dec. 31, 2019	Oct. 31, 2019	Dec. 31, 2021	Dec. 31, 2020	Dec. 31, 2019	Nov. 30, 2020
Related Party Transaction [Line Items]
Shares issued as payment for services								$ 10,446
Proceeds from sales of equity securities					$ 21,587	$ 0	$ 0	23,456
Deferred revenue		$ 25,823				27,465	25,823
Exotech, Inc.; AD Skincare, Inc.; and Thrive Agrobiotics, Inc.
Related Party Transaction [Line Items]
Investments in affiliates									$ 326
Collaboration and licensing agreements
Related Party Transaction [Line Items]
Deferred revenue		$ 23,420				23,023	23,420
Harvest Intrexon Enterprise Fund I, LP
Related Party Transaction [Line Items]
Stock issued in conjunction with settlement agreement, shares		2,117,264
Shares issued in conjunction with settlement agreement		$ 18,103
Loss related to litigation settlement		$ 11,436
Exotech, Inc.; AD Skincare, Inc.; and Thrive Agrobiotics, Inc. | Collaboration and licensing agreements
Related Party Transaction [Line Items]
Deferred revenue									$ 6,993
Third Security
Related Party Transaction [Line Items]
Expense for services						100	159	$ 26
Common stock calculation period								30 days
Shares issued as payments for services (in shares)								1,606,062
Shares issued as payment for services								$ 8,233
Sublease rental income						$ 75	$ 83	89
Cash received on contract termination	$ 143
Castle Creek Biosciences, Inc.
Related Party Transaction [Line Items]
Proceeds from sales of equity securities				$ 1,280
Proceeds from convertible preferred shares and convertible note			$ 3,311
Changes in fair value, gain								3,222
Common Stock | Third Security
Related Party Transaction [Line Items]
Expense for services								$ 800
Third Security | Executive Chairman
Related Party Transaction [Line Items]
Ownership interest						100.00%